Haplogroup U4 is a small Indo-European haplogroup.
Here are sequences for this haplogroup:
1. Annunen-Rasila AM260586 CODING AREA ONLY
2. Annunen-Rasila AM260587 CODING AREA ONLY
3. Moilanen (3)A20.JS AY339549(Finland)
4. Moilanen (3)A21.JS AY339550(Finland)
5. Achilli_Rengo A8.JS AY882386(Spain)
6 Achilli_Rengo A9.JS AY882387(Italy)
7. Achilli_Rengo A10.JS AY882388(Adygei)
8. Kivisild (2)A7.JS DQ112762(N. Europe) CODING AREA ONLY
9. Kivisild (3)A34.JS DQ112859(Asia) CODING AREA ONLY
10. Herrn.-Europe (2)A35.JS mtDNA109(Europe) CODING AREA ONLY
11. Herrn.-Europe (3)A57.JS mtDNA272(Europe) CODING AREA ONLY
12. Herrn.-Europe (4)A62.JS mtDNA359(Europe) CODING AREA ONLY
13. Herrn.-Europe (6)A31.JS mtDNA502(Europe) CODING AREA ONLY
14. Herrn.-Europe (6)A45.JS mtDNA517(Europe) CODING AREA ONLY
15. Montiel-Sosa A2.JS DQ156209
16 FTDNA A32.JS EF452295
17. Gasparre1 A10.JS EF660921(Italy)
18. Gasparre2 A3.JS EF660965(Italy)
19. Gasparre2 A21.JS EF660983(Italy)
20. La Morgia A2.JS EF060364
21. Grzybowski EF222235
22. Grzybowski EF222236
23. Grzybowski EF222237
24. Grzybowski EF222238
25. Grzybowski EF222239
26. Grzybowski EF222240
27. Grzybowski EF222241
28. Grzybowski EF222242
29. Grzybowski EF222248
30. Grzybowski EF222249
31. Grzybowski EF222250
32. Grzybowski EF222252
33. FTDNA EF452295
34. Ingman_gyll EU007874(Kchanti)
35. Ingman_gyll EU007891(Mongolia)
36. FTDNA EU135973
37. FTDNA EU140898
38. FTDNA EU428753
39. Hinttala DQ489514 CODING AREA ONLY
40. Maasz EU571946(Hungary)
The sequences have these differences from the CRS:
- for the main stem leading from H:
A73G A263G A750G A1438G A2706G A4769G
C7028T A8860G G11719A C14766T A15326G
and for Haplogroup U
A11467G A12308G G12372A
for U4 & U9 (U9 also - 3531 3584 6386 14094)
T152C(varies) G499A A1811G T5999C
and for U4
T195C T4646C A6047G C11332T C14620T T15693C T16356C(varies)
- excluding changes at 309 & 315
- and changes around 523
- variation of T16519C
Subgroups:
U4a - C8818T
U4b - T7705C
U4c - A4811G
The mutation lists are:
1. Annunen-Rasila AM260586 (15693) CODING AREA ONLY U4d
T629C 2405.C C2772T A5319T A5984G C6938T T8260C
2. Annunen-Rasila AM260587 (15693) CODING AREA ONLY U4d
T629C 2405.C C2772T A5319T A5984G C6938T T8260C
3. Moilanen (3)A20.JS AY339549(Finland) (11332) U4d
2405.C C2772T A5984G C6653A C6938T T8260C
4. Moilanen (3)A21.JS AY339550(Finland) U4a
T310C C8818T G15497A C16223T
5. Achilli_Rengo A8.JS AY882386(Spain) U4a
T961C 965.C 965.C 965.C A1555G A4562G C8818T A12937G C16134T
6. Achilli_Rengo A9.JS AY882387(Italy) U4a
T310C A1811G C8818T T16359C
7. Achilli_Rengo A10.JS AY882388(Adygei) U4b
T2083C A3672G T7705C A8642G T11339C T12297C C15789T T16362C
8. Kivisild (2)A7.JS DQ112762(N. Europe) U4c
A4811G A6146G 8276.C 8276.C 8276.C 8276.C T9070G T10907C T11009C C14866T
9. Kivisild (3)A34.JS DQ112859(Asia) U4a
G5460A C6164T C8818T T11204C C12888T T13899C
10. Herrn.-Europe (2)A35.JS EF657242 mtDNA109(Europe) U4b
T7705C T11339C C11932T A13528G C13565T
11. Herrn.-Europe (3)A57.JS EF657423 mtDNA272(Europe) U4c
T2351C A4811G A6146G T9070G T10907C A13973T C14866T
12. Herrn.-Europe (4)A62.JS EF657519 mtDNA359(Europe) U4c
A2905G C3654T A4811G A6146G T9070G T10907C C14866T
13. Herrn.-Europe (6)A31.JS EF657679 mtDNA502(Europe) U4b
A3672G T7705C C11143T G11176A T11339C T12609C G13708A
14. Herrn.-Europe (6)A45.JS EF657695 mtDNA517(Europe) U4b
T7705C T11339C A13528G C13565T
15. Montiel-Sosa A2.JS DQ156209 U4a
T310C C8818T
16. FTDNA A32.JS EF452295 U4b
C1185T T7705C T8433C T14178C G15883A T16136C C16278T
17. Gasparre1 A10.JS EF660921(Italy) U4b
A3672G T7705C C11143T G11176A T11339C T12609C G13708A C15632T T16311C
18. Gasparre2 A3.JS EF660965(Italy) U4a
A153G T961C 965.C 965.C T7084C C8818T A12937G C16134T T16144C
19. Gasparre2 A21.JS EF660983(Italy) U4b
T2083C A3672G T7705C A8642G T11339C T12297C C13272T G15140A C15789T T16362C
20. La Morgia A2.JS EF060364 U4a
G247A A801G A2792G G3460A C6665T A6929G G8065A C8818T G15773A T16362C
21. Grzybowski EF222235 U4a
T310C G5261A C8818T
22. Grzybowski EF222236 U4a
T310C T8496C C8818T
23. Grzybowski EF222237 U4a
T146C T310C A6800G T8496C C8818T
24. Grzybowski EF222238 U4a
T310C C8818T G13759A T16362C
25. Grzybowski EF222239 U4a
T310C G8557A C8818T T9311C T14319C C16294T
26. Grzybowski EF222240 U4a
T310C C8818T G9948A T16172C C16223T
27. Grzybowski EF222241 U4a
T310C T2887C C8818T C16223T
28. Grzybowski EF222242 U4a
T310C T7051C T8567C C8818T C10654T C16242A G16274A T16288C T16362C
29. Grzybowski EF222248 U4c
A4811G A6146G T9070G G9133A T10907C T11009C C14866T C16179T
30. Grzybowski EF222249 U4a
T310C C8818T
31. Grzybowski EF222250 U4a
T310C T8496C C8818T T16189C
32. Grzybowski EF222252 U4a
T310C T8567C C8818T C10654T C16242A T16288C T16362C
33. FTDNA EF452295 U4b
C1185T T7705C T8433C T14178C G15883A T16136C C16278T
34. Ingman_gyll EU007874(Kchanti) U4b
A215G C3204T A3434G T7705C
35. Ingman_gyll EU007891(Mongolia) U4d
T629C 2405.1C T5567C C10692T C11326T G11518A A13105G
36. FTDNA EU135973 U4c
C150T A4811G A6146G T9070G T10907C T11009C C14866T C16179T
37. FTDNA EU140898 U4b
T7705C T16136C A16233G
38. FTDNA EU428753 U4a
C296T A745G C3204T G7119A T8705C C8818T A12937G C14668T G16129A C16134T
39. Hinttala DQ489514 CODING AREA ONLY U4a
A2792G C6665T A6929G G7497A G8065A T8227C C8818T A12924G
40. Maasz EU571946(Hungary) U4b
T146C A3672G A5752. A7265G A7445G T7705C T9989C T11339C C11674T C15481T
41. Grzybowski A23.JS EU545415(Belarus) Haplogroup U4b
A73G T195C A263G G499A A750G A1438G A1811G A2706G C3107N A3672G
T4646C A4769G T5999C A6047G C7028T T7705C A8860G C11143T G11176A C11332T
T11339C A11467G G11719A A12308G G12372A T12609C G13708A C14620T C14766T A15326G
C15632T T15693C T16356C T16519C
42. Grzybowski A24.JS EU545416(Poland) Haplogroup U4a1
A73G T152C T195C T204C A263G G499A A750G T961C A1438G A1811G
A2706G C3107N T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T
A11467G G11719A A12308G G12372A A12937G C14620T C14766T A15326G T15693C C16134T
A16207G T16356C T16519C
43. Grzybowski A25.JS EU545417(Poland) Haplogroup U4a1
A73G T152C T195C A263G G499A A750G T961C A1438G A1811G A2706G
C3107N T4646C A4769G T5999C A6047G C7028T A7271G C8818T A8860G C11332T
A11467G G11719A A12308G G12372A A12937G C14620T C14766T C14884T A15326G T15693C
C16134T T16356C T16519C
44. Grzybowski A26.JS EU545418(Poland) Haplogroup U4a1
A73G T152C T195C A263G C296T G499A A745G A750G A1438G A1811G
A2706G C3107N C3204T T4646C A4769G T5999C A6047G C7028T G7853A C8818T
A8860G C11332T A11467G G11719A A12308G G12372A A12937G C14368T C14620T C14766T
A15326G T15693C C16134T T16356C T16519C
45. Grzybowski A27.JS EU545419(Poland) Haplogroup U4b
A73G T146C A263G G499A A750G A1438G A1811G A2706G C3107N A3672G
T4646C A4769G T5999C A6047G C7028T T7705C A8860G T9989C C11332T T11339C
A11467G C11674T G11719A A12308G G12372A C14620T C14766T A15326G T15693C T16356C
T16519C
46. Grzybowski A29.JS EU545421(Czech) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G G11719A
A12308G G12372A C14620T C14766T A15326G T15693C C16295T T16356C T16519C
47. Grzybowski A31.JS EU545423(Czech) Haplogroup U4a3
A73G T195C G247A A263G G499A A750G A1438G A1811G A2706G A2792G
C3107N T4646C A4769G T5999C A6047G C6665T A6929G C7028T G8065A C8818T
A8860G C11332T A11467G G11719A A12308G G12372A C14620T C14766T A15326G T15693C
A16265G T16356C T16362C T16519C
48. Grzybowski A33.JS EU545425(Czech) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G G11719A
C11881T A12308G G12372A C14620T C14766T A15326G T15693C T16519C
49. Grzybowski A34.JS EU545426(Czech) Haplogroup U4a2
A73G T152C G185A A189G T195C A215G A263G G499A T629C A750G
T1193C A1438G A1811G A2706G C3107N T4646C A4769G T5567C T5999C A6047G
C7028T A8860G C10692T C11326T C11332T A11467G G11518A G11719A A12308G G12372A
A13105G G13145A C14620T C14766T A15326G T15693C T16356C T16519C
50. Grzybowski A35.JS EU545427(Czech) Haplogroup U4a1
A73G T152C T195C A263G G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T G8155A C8818T A8860G C11332T A11467G
G11719A A12308G G12372A A12937G A13158G T14110C C14620T C14766T A15326G T15693C
C16134T C16221T C16234T T16356C T16519C
51. Grzybowski A36.JS EU545428(Poland) Haplogroup U4a1
A73G T152C T195C A263G G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T G8155A C8818T A8860G C11332T A11467G
G11719A A12308G G12372A A12937G A13158G T14110C C14620T C14766T A15326G T15693C
C16134T C16234T T16356C T16519C
52. Grzybowski A37.JS EU545429(Poland) Haplogroup U4a1
A73G T152C T195C A263G G499A A745G A750G A1438G A1811G A2706G
C3107N C3204T T4646C A4769G T5999C A6047G C7028T C7816T C8818T A8860G
C11332T A11467G G11719A T11809C A12308G G12372A A12937G C14620T C14766T A15326G
T15693C C16134T C16301T T16356C T16519C
53. Grzybowski A38.JS EU545430(Belarus) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G G11719A
A12308G G12372A C14620T C14766T A15326G T15693C T16356C T16519C
54. Grzybowski A39.JS EU545431(Belarus) Haplogroup U4a1
A73G T152C T195C A263G G499A A750G T961C A1438G A1811G A2706G
C3107N T4646C A4769G T5999C A6047G C7028T T8167C C8818T A8860G C11332T
A11467G G11719A A12308G G12372A G12618A A12937G C14620T C14766T A15326G T15693C
C16134T T16356C T16519C
55. Grzybowski A40.JS EU545432(Poland) Haplogroup U4a1
A73G T195C A263G G499A A745G A750G A1438G A1811G A2706G C3107N
C3204T T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G
G11719A A12308G G12372A A12937G C14568T C14620T C14766T A15326G T15693C C16134T
C16301T T16356C T16519C
56. Grzybowski A41.JS EU545433(Poland) Haplogroup U4a1
A73G T152C T195C A263G G499A A750G T961C A1438G A1811G A2706G
C3107N T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G
G11719A A12308G G12372A A12937G C14620T C14766T A15326G T15693C C16134T T16189C
T16356C T16519C
57. Grzybowski A42.JS EU545434(Russia) Haplogroup U4a2
A73G G143A T195C A214G A263G T310C G499A A750G A1438G A1811G
A2706G C3107N T4646C A4769G T5999C A6047G C6164T C7028T C8818T A8860G
C11332T A11467G G11719A A12308G G12372A T12624C C14620T C14766T A15326G T15693C
T16356C T16519C
58. Grzybowski A46.JS EU545438(Russia) Haplogroup U4d
A73G T195C A263G G499A T629C A750G A1438G A1811G A2706G C2772T
C3107N T4646C A4769G T5999C A6047G C7028T A8860G C11332T A11467G G11719A
A12308G G12372A A13398G C14620T C14766T A15326G T15693C T16356C T16519C
59. Grzybowski A48.JS EU545440(Russia) Haplogroup U4a2
A73G T152C T195C A263G T310C G499A A750G A1438G A1811G A2706G
C3107N T4646C A4769G T5999C A6047G C7028T A7424G C8818T A8860G C11332T
A11467G G11719A A12308G G12372A C14620T C14766T A15326G T15693C C16150T T16189C
C16360T T16519C
60. Grzybowski A50.JS EU545442(Russia) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2246G A2706G
C3107N T4646C A4769G T5999C A6047G G6383A C7028T C8818T A8860G C11332T
A11467G G11719A A12308G G12372A C14620T A14696G C14766T A15326G T15693C T16356C
T16519C
61. Grzybowski A52.JS EU545444(Russia) Haplogroup U4b
A73G T195C A263G G499A A750G A1438G A1811G A2706G C3107N T4646C
A4769G T5999C A6047G C7028T T7705C A8860G A10523G C11332T A11467G G11719A
A12308G G12372A C14620T C14766T A15326G T15693C T16356C T16519C
62. Grzybowski A54.JS EU545446(Russia) Haplogroup U4a2
A73G T195C A263G T310C G499A G709A A750G A1438G A1811G A2706G
C3107N T4646C A4769G C5945T T5999C A6047G C7028T C8818T A8860G C11332T
A11467G G11719A A12308G G12372A T14319C C14620T C14766T A15326G T15693C C16294T
T16519C
63. Grzybowski A56.JS EU545448(Russia) Haplogroup U4d
A73G T195C A263G G499A T629C A750G A1438G A1811G A2706G C2772T
C3107N T4646C A4769G T5999C A6047G C7028T A8860G C11332T A11467G G11719A
A12308G G12372A A13398G C14620T C14766T A15326G T15693C T16356C T16519C
64. Grzybowski A58.JS EU545450(Russia) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G G11719A
G12192A A12308G G12372A C14620T C14766T A15326G T15693C C16169T G16319A T16519C
65. Grzybowski A59.JS EU545451(Russia) Haplogroup U4a1
A73G T152C T195C A263G G499A A750G T961C A1438G A1811G A2706G
C3107N T4646C A4769G T5999C A6047G C7028T T8167C C8818T A8860G T10790C
C11332T A11467G G11719A A12308G G12372A G12618A A12937G C14620T C14766T A15326G
T15693C C16134T T16356C T16519C
66. Grzybowski A61.JS EU545453(Slovakia) Haplogroup U4a2
Leu(CUN).. A12308G
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T G8027A C8818T A8860G C11332T A11467G
G11719A A12021C A12308G G12372A C14620T C14766T A15326G T15693C T16356C T16519C
67. Grzybowski A63.JS EU545455(Slovakia) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G G11719A
A12308G G12372A C14620T C14766T A15326G T15693C A16183C T16189C T16356C T16519C
68. Grzybowski A64.JS EU545456(Slovakia) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2246G A2706G
C3107N T4646C A4769G T5999C A6047G G6383A C7028T C8818T A8860G C11332T
A11467G G11719A A12308G G12372A C14620T A14696G C14766T A15326G T15693C T16356C
T16519C
69. Grzybowski A66.JS EU545458(Slovakia) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G G11719A
A12308G G12372A C14620T C14766T A15326G T15693C T16519C
70. Grzybowski A67.JS EU545459(Slovakia) Haplogroup U4a1
A73G T152C T195C A263G C296T G499A A745G A750G A1438G A1811G
A2706G C3107N C3204T T4646C A4769G T5999C A6047G C7028T C8818T A8860G
C11332T A11467G G11719A A12308G G12372A A12937G C14620T C14766T A15326G T15693C
C16134T T16356C T16519C
71. Grzybowski A68.JS EU545460(Slovakia) Haplogroup U4a2
A73G T195C A263G G499A A750G A1438G A1811G A2706G C3107N T4646C
A4769G T5999C A6047G C7028T G7762A T8705C A8860G T10118C C10676T T10907C
C11332T A11467G C11497T G11719A A12308G G12372A C14620T C14766T A15326G T15693C
C16261T T16356C
72. Grzybowski A69.JS EU545461(Slovakia) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T C8818T A8860G C11332T A11467G G11719A
A12308G G12372A G13759A C14620T C14766T A15326G T15693C T16519C
73. Grzybowski A70.JS EU545462(Slovakia) Haplogroup U4c
A73G T195C A263G G499A A750G A1438G C1788T A1811G A2706G C3107N
T4646C A4769G A4811G T5999C A6047G A6146G C7028T A8860G T9070G T10907C
T11009C C11332T A11467G G11719A A12182G A12308G G12372A C14620T C14766T C14866T
A15326G T15693C C16179T T16356C T16519C
74. Grzybowski A71.JS EU545463(Slovakia) Haplogroup U4b
A73G T146C T152C T195C A263G G499A A750G T1193C A1438G A1811G
A2706G C3107N T4646C A4769G T5999C A6047G C7028T T7705C A8860G C11332T
T11339C A11467G G11719A A12308G G12372A A13528G C13565T C14620T C14766T A15326G
T15693C T16356C T16519C
75. Grzybowski A72.JS EU545464(Belarus) Haplogroup U4a2
A73G T195C A263G T310C G499A A750G A1438G A1811G A2706G C3107N
T4646C A4769G T5999C A6047G C7028T T8567C C8818T A8860G G9548A C11332T
A11467G G11719A A12308G G12372A C14620T A15326G T15693C T16356C T16519C
76. Grzybowski A73.JS EU545465(Belarus) Haplogroup U4c
A73G T195C A263G G499A A750G A1438G A1811G A2706G C3107N T4646C
A4769G A4811G T5999C A6047G A6146G C7028T A8860G T9070G T10907C T11009C
C11332T A11467G G11719A A12308G G12372A C14620T C14766T C14866T A15326G T15693C
C16179T T16356C T16519C